AN exceptionally preserved new specimen of the giant conodont Promissum pulchrum reveals details of the trunk musculature, feeding apparatus and eyes, High-fidelity resolution of ultrastructural features of the trunk myomeres provides the first conclusive evidence of muscle-fibre organization and orientation in an extinct agnathan. The presence of fibrous extrinsic eye muscles confirms the degree of encephalization of the conodonts and is consistent with a cladistic position crownwards of the myxinoids. The soft tissues are uniquely preserved as illite and mixed-layer clay minerals.